A live but retracted individual of Laemodonta siamensis on a human hand. A partially torn epiphragm of dried mucus is visible in the aperture of the shell
In this family, the number of haploidchromosomes lies between 16 and 20 (according to the values in this table).[4]
They have a distinctive mode of locomotion arising due to a split sole. The front part of the sole adheres to the substrate and then the rear part of the sole is drawn up to the front part.[5]
Taxonomy
Species are traditionally classified into five taxonomic groups, the Pythiidae, the Laemodontidae, the Melampodidae, the Ellobiidae and the Carychiidae.[1] These taxa have been recognized either as families within Ellobioidea or as sub-families within the family Ellobiidae.[1]
The high degree of homoplasy in morphological characters and frequent low variability has led to the description of approximately 800 species names available in the literature, of which 250 are likely to be valid.[1]
A molecular study by Dayrat at al. (2011)[6] for the Ellobioidea suggests a monophyletic origin of the entire group.[1]
† Zaptychius Walcott, 1883 - type genus of the subfamily
Genera brought into synonymy
Alexia Leach in Gray, 1847: synonym of Myosotella Monterosato, 1906
Apodosis Pilsbry & McGinty, 1949 synonym of Leuconopsis F. W. Hutton, 1883 (junior subjective synonym)
Auricula Lamarck, 1799: synonym of Ellobium Röding, 1798
Auriculodes Strand, 1928: synonym of Ellobium (Auriculodes) Strand, 1928 represented as Ellobium Röding, 1798
Auriculus Montfort, 1810 synonym of Ellobium Röding, 1798 (unjustified emendation of Auricula)
Autonoe Guppy, 1868 synonym of Ellobium (Auriculodes) Strand, 1928 represented as Ellobium Röding, 1798 (junior homonym of Autonoe Leach, 1852 [Bivalvia]; Autonoella Wenz, 1947, is a replacement name)
Cassidulus Beck, 1838 synonym of Cassidula A. Férussac, 1821 (Invalid: unjustified emendation of Cassidula Férussac, 1821, and also junior homonym of Cassidulus Lamarck, 1801 [Echinodermata])
Cremnobates Swainson, 1855: synonym of Marinula King & Broderip, 1832
Cylindrotis Möllendorff, 1895 synonym of Auriculastra E. von Martens, 1880
DetraciaGray in Turton, 1840: synonym of Melampus (Detracia) Gray, 1840 represented as Melampus Montfort, 1810
Jaminia T. Brown, 1827: synonym of Myosotella Monterosato, 1906
Kochia Pallary, 1900: synonym of Myosotella Monterosato, 1906
Laimodonta H. Adams & A. Adams, 1855: synonym of Laemodonta Philippi, 1846
Leuconia Gray, 1840: synonym of Auriculinella Tausch, 1886
Leucopepla Peile, 1926: synonym of Auriculinella Tausch, 1886
Leucophytia Winckworth, 1949: synonym of Auriculinella Tausch, 1886
Maripythia Iredale, 1936: synonym of Marinula P. P. King, 1832 (unavailable name: no description)
Melosidula B. J. Smith & Kershaw, 1979: synonym of Cassidula (Cassidula) A. Férussac, 1821 represented as Cassidula A. Férussac, 1821 (junior subjective synonym)
Monica H. Adams & A. Adams, 1855: synonym of Ovatella Bivona-Bernardi, 1832
Nealexia Wenz, 1920: synonym of Myosotella Monterosato, 1906
Pira H. & A. Adams, 1855: synonym of Melampus Montfort, 1810
Plecotrema H. Adams & A. Adams, 1854: synonym of Laemodonta Philippi, 1846
Rangitotoa Powell, 1933: synonym of Microtralia Dall, 1894
Rhodostoma Swainson, 1840: synonym of Cassidula A. Férussac, 1821 (junior subjective synonym)
Sidula J. E. Gray, 1840: synonym of Cassidula A. Férussac, 1821 (junior subjective synonym)
Tifata H. Adams & A. Adams, 1855: synonym of Melampus (Detracia) Gray, 1840 represented as Melampus Montfort, 1810 (junior subjective synonym)
Volatela Starobogatov, 1993: synonym of Myosotella Monterosato, 1906
Taxonomic note
Genus Sarnia H. Adams & A. Adams, 1855 (temporary name, Sarnia [H. & A. Adams, 1855 (September), The Genera of Recent Mollusca, 2: 239] is an incorrect subsequent spelling of Siona H. Adams & A. Adams, 1855 which is a junior homonym of Siona Duponchel, 1829 [Lepidoptera].)
Ecology
The taxon Ellobioidea comprises a group of morphologically and ecologically highly diverse snails, known to have successfully invaded the marine, brackish water and terrestrial habitats.[1]
These are mostly snails that live in salt marshes and similar maritime habitats, and thus have a tolerance for saline conditions.
References
123456Weigand A. M., Jochum A., Slapnik R., Schnitzler J., Zarza E. & Klussmann-Kolb A. (2013). "Evolution of microgastropods (Ellobioidea, Carychiidae): integrating taxonomic, phylogenetic and evolutionary hypotheses". BMC Evolutionary Biology13: 18. doi:10.1186/1471-2148-13-18This article incorporates textfrom this source, which is available under the CC BY 2.0 license.
↑Barker G. M.: Gastropods on Land: Phylogeny, Diversity and Adaptive Morphology. in Barker G. M. (ed.): The biology of terrestrial molluscs. CABI Publishing, Oxon, UK, 2001, ISBN0-85199-318-4. 1-146, cited pages: 139 and 142.
↑Herbert,dai and, Kilburn, Dick (2004). Field Guide to the Land Snails and Slugs of Eastern South Africa. Scottsville, South Africa: Natal Museum. p.98. ISBN0-620-32415-5.{{cite book}}: CS1 maint: multiple names: authors list (link)
↑Dayrat B., Conrad M., Balayan S., White T. R., Albrecht C., Golding R., Gomes S., Harasewych M. G., de Frias Martins A. M. (2011). "Phylogenetic relationships and evolution of pulmonate gastropods (Mollusca): new insights from increased taxon sampling". Molecular Phylogenetics and Evolution59: 425-437. doi:10.1016/j.ympev.2011.02.014.