Capulus Temporal range: 443.8–0.0 Ma PreꞒ Ꞓ O S D C P T J K Pg N Silurian to Recent
Two shells of Capulus ungaricus with the periostracum removed
Scientific classification
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Subclass:	Caenogastropoda
Order:	Littorinimorpha
Superfamily:	Capuloidea
Family:	Capulidae
Genus:	Capulus Montfort, 1810
Type species
Patella ungarica Linnaeus, 1758
Synonyms
Actita Fischer von Waldheim, 1823 (Unnecessary substitute name for Capulus and Pileopsis) Brocchia Bronn, 1828 Capulonix Iredale, 1929 Pileopsis Lamarck, 1822 Pileopsis (Brocchia) Bronn, 1828

Capulus is a genus of small sea snails, marine gastropod mollusks in the family Capulidae, the cap snails.^[1]

Species

Species within the genus Capulus include:

• Capulus badius Dunker, 1882
• Capulus californicus Dall, 1900
• Capulus compressus E. A. Smith, 1891
• Capulus daanbantayanensis Poppe & Tagaro, 2026
• Capulus danieli (Crosse, 1858)
• † Capulus deurganckensis Marquet & Landau, 2006
• Capulus devexus (May, 1916)
• Capulus dilatatus A. Adams, 1860
• Capulus elegans (Tapparone Canefri, 1877)
• Capulus fragilis E. A. Smith, 1904
• Capulus huangi S.-I Huang & Y.-F. Huang, 2012
• Capulus japonicus A. Adams, 1861
• Capulus kawamurai Habe, 1992
• Capulus ngai Thach, 2016
• Capulus novaezelandiae Dell, 1978
• Capulus otohimeae (Habe, 1946)
• † Capulus partimsinuosus (S. V. Wood, 1848)
• Capulus sericeus J. Burch & R. Burch, 1961
• Capulus simplex Locard, 1898^[2]
• † Capulus sinuosus (Brocchi, 1814)
• Capulus spondylicola Habe, 1967
• Capulus subcompressus Pelseneer, 1903^[3]
• Capulus ungaricoides (d'Orbigny, 1841)
• Capulus ungaricus (Linnaeus, 1758)^[4]
• Capulus violaceus Angas, 1867

Species brought into synonymy

• Capulus bicarinatus (Pease, 1861a): synonym of Amathina bicarinata Pease, 1861
• Capulus chilensis Dall, 1904: synonym of Capulus ungaricoides (d'Orbigny, 1841)
• † Capulus corrugatus (extinct/fossil):^[5] synonym of † Capulus partimsinuosus (S. V. Wood, 1848)
• Capulus devotus Hedley, 1904: synonym of Malluvium devotum (Hedley, 1904) (original combination)
• Capulus galea Dall, 1889: synonym of Hyalorisia galea (Dall, 1889)
• Capulus galeus [sic]: synonym of Capulus galea Dall, 1889: synonym of Hyalorisia galea (Dall, 1889)
• Capulus hungaricus [sic]: synonym of Capulus ungaricus (Linnaeus, 1758)
• Capulus incurvus (Gmelin, 1791): synonym of Hipponix incurvus (Gmelin, 1791)
• Capulus intortus (Lamarck, 1822):^[6] synonym of Hipponix incurvus (Gmelin, 1791)
• Capulus kawamurai Habe, 1992: synonym of Capulus danieli (Crosse, 1858)
• Capulus liberatus Pease, 1868: synonym of Krebsia liberata (Pease, 1868)
• Capulus lissus E.A. Smith, 1894: synonym of Malluvium lissum (E. A. Smith, 1894)
• Capulus militaris (Linnaeus, 1771): synonym of Capulus ungaricus (Linnaeus, 1758)
• Capulus nutatus Hedley, 1908: synonym of Williamia radiata nutata (Hedley, 1908)
• Capulus radiatus M. Sars, 1851: synonym of Piliscus radiatus (Sars M., 1851): synonym of Piliscus commodus (Middendorff, 1851)
• Capulus sagittifer Gould, 1852: synonym of Phenacolepas sagittifer (Gould, 1852)
• Capulus shreevei Conrad, 1869: synonym of Cyrtopleura costata (Linnaeus, 1758)
• Capulus sycophanta Garrard, 1961: synonym of Capulus danieli (Crosse, 1858)
• Capulus uncinatus (Hutton, 1873): synonym of Capulus danieli (Crosse, 1858)

According to the Indo-Pacific Molluscan Database (OBIS) the following species names are also in current use ^[7]

• Capulus irregularis E. A. Smith, 1895
• Capulus paleacea Menke

References

• Beu A.G. (2004). "Marine Mollusca of Oxygen Isotope Stages of the Last 2 Million Years in New Zealand. Part 1: Revised Generic Positions and Recognition of Warm-Water and Cool-Water Migrants". Journal of the Royal Society of New Zealand 34(2): 192.
• Gofas, S.; Le Renard, J.; Bouchet, P. (2001). "Mollusca". In: Costello, M.J. et al. (ed.) (2001). European Register of Marine Species: A Check-List of the Marine Species in Europe and a Bibliography of Guides to Their Identification. Collection Patrimoines Naturels, 50: pp. 180–213.
• Spencer, H.; Marshall. B. (2009). "All Mollusca Except Opisthobranchia". In: Gordon, D. (ed.) (2009). New Zealand Inventory of Biodiversity. Volume One: Kingdom Animalia. 584 pp.

External links

Wikimedia Commons has media related to Capulus.

• Montfort P. [Denys de]. (1808-1810). Conchyliologie systématique et classification méthodique des coquilles. Paris: Schoell. Vol. 1: pp. lxxxvii + 409 [1808]. Vol. 2: pp. 676 + 16